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DISTRIBUTION OF SUCCULENTS
The highest concentration of
succulents
is found at a latitude between 40° north and 40° South of the Equator.
Some, however, may also occur as far north as the 56th parallel while the
southernmost location is at about 50° south latitude. This large and
geomorphologically diverse belt consists mainly of arid lands characterised
by long dry spells interspersed with occasional rainy periods or desert
regions where fog and light drizzle are quite common, like the coastal strip
of Namibia and the plateau lying between the Andes and the pacific coast in
South America but it also includes humid and hot tropical rainforests,
steppes, savannahs, highlands and mountain ranges. A few species or genera,
thanks to their vigour and adaptability, are able to colonize vast
territories, as is the case with the widespread genus Opuntia; others are
found only in more restricted areas. Humans played a fundamental part in the
diffusion of certain species across different continents, helped by the fact
that these extraordinary plants can often cope with adverse conditions.
By and large, the different habitats can be divided into the following four
categories:
- pseudo-deserts and subdeserts;
- steppes and prairies;
- mountainous land;
- tropical and subtropical forests.
The first environment is distinguished by prolonged droughts punctuated with
erratic heavy downpours (total rainfall between 500 and 800 mm. per year);
regions receiving less than 250 mm. of rain per year are true deserts, such
as the Sahara, that are inhospitable and cannot sustain the life of plants,
not even succulents. This is not quite true when it comes to deserts like
those of Namibia, Chile and Peru where the supply of air moisture ensured by
the frequent fogs compensates for the scarcity of rain. Many genera of
cactus live here: Oroya, Borzicactus, Chamaecereus, Eriosyce, Eulychnia,
Haageocereus, Neoraimondia, Oreocereus, Rebutia, just to mention a few.
Another habitat is found in the vast plains and highlands like those of
Mexico and southwestern US (Texas, New Mexico, Arizona), where Cactaceae
like Mammillaria and Echinocereus generally predominate; some plants, for
example Aylostera, Echinofossulocactus, Weingarthia, grow on the bare rock
of mountain slopes. This area is characterized by great diurnal and annual
temperature variation and here snow may also occur. The Central States of
Mexico are home to a great variety of species, among which Ariocarpus,
Atzekium, Pelecyphora and Obregonia are the most prized by collectors. The
well-known Cephalocereus senilis grow in the State of Hidalgo while
Lophocereus schottii and Machaerocereus eruca are native to Baja California;
Pachycereus and Lemaireocereus may be found in other regions.
As we proceed towards the Equator we encounter the equatorial
rainforests, typical of Central America, Guyana, the Amazon region and the
West Indies, where cacti grow on trees as epiphytic plants, yet not
parasitic, developing their roots in the leaf litter that accumulates on
horizontal branches or at the bifurcation of trunks. Here species like
Selenicereus, Hylocereus, Epiphyllum, Hoya, Rhipsalis and Schlumbergera
thrive, growing aerial roots that can absorb moisture from the humid air
when there is lack of rain. Melocactus are native to the West Indies and
require warm temperatures throughout the year.
Moving further south towards Venezuela, Equador, Peru, Bolivia, Chile
and western regions of Northern Argentina we come across many genera that
inhabit North America as well, though represented by different species; here
Cerei are predominant, particularly those with dense hairs like Espostoa and
Oreocereus. Matucana, Mila, Oroya grow on the high-altitude plains. In the
Chilean Andes and the Atacama desert of Chile we can find genera like
Browningia, Eulychinia, Neoporteria, Copiapoa. The Andinian plants are known
to be quite difficult in cultivation, especially the ones from Peruvian and
Chilean mountains which grow at great elevation under high light levels;
yet, the almost constant cloud cover prevents their epidermis from scorching
while if grown where sunshine amounts are greater, even at higher latitudes,
they can easily get burned. Steppes and prairies are home to many genera
that are well-known to cactus enthusiasts such as Cleistocactus, Notocactus,
Gymnocalycium, Trichocereus, Parodia, Rebutia, Lobivia and Echinopsis. On
Brazilian highlands we encounter Cereus, Harrisia, Trichocereus; in Southern
Brazil as well as Paraguay and Uruguay again we might come across many Cerei
while epiphytic cacti inhabit coastal forests. The Pampas is the natural
habitat of Trichocereus, Harrisia, Cleistocactus, Chamaecereus, Echinopsis,
Gymnocalycium, Notocactus.
Generally speaking, we can say that
Cactaceae come from
the Americas in the same way that the majority of other
succulents comes from
Africa although the latter may also occur in Europe, Asia and Australia.
Some genera like Aloe, Gasteria and Haworthia are unique to Africa,
similarly to Pelargoniums though a few species of this genus can also be
found outside the continent. Most succulents and cacti are concentrated
mainly across the tropical and subtropical belts throughout the world but
they occur in many widely scattered places so no real attempt has ever been
made to give an exhaustive description of all these different habitats.
Here we find Euphorbias, growing almost over the entire area thanks to
the great diversity of species. The range of the Asclepiadaceae family
stretches from the Himalayan mountain range to the Far East, reaching as far
as Australia although some genera are unique to Africa, as is the case with
Huernia, Hoodia, Ceropegia, Caralluma and Stapelia. Crassulaceae grow in
South Africa, Namibia, Madagascar but also in the U.S., Mexico as well as
Europe (particularly Sedum and Sempervivum). The various species of Agave
are native to the Americas, along with Yucca, Echeveria and Duddleya. Mexico
is home to Pachyphytum, Sedum and Idria columnaris, among others. Epiphytic
and non-epiphytic Bromeliaceae inhabit humid and warm places of the American
continents. Eastern Africa is the habitat of many species of Euphorbia, Aloe
and Stapelia; some Euphorbias are also found in Madagascar together with
Kalanchoe. Monanthes and Aeonium are from the Canary Islands.
On the whole we can say that Anacardiaceae occur in the tropical and
subtropical belts of the northern hemisphere, Apocynaceae, Burseraceae,
Commelinaceae, Rubiaceae and Sterculiaceae mainly in the tropics,
Bombacaceae in tropical and subtropical regions of Africa, Madagascar, the
Indian Ocean and Australia and finally Compositae, Convolvulaceae,
Liliaceae, Oxalidaceae, Passifloraceae, Vitaceae, Pedaliaceae mostly in
Africa.
The South African Veld, with its great concentration and variety of
succulent families and species is the paradise of the Mesembryanthemaceae.
These plants occur from sea level to 3000 metres of altitude (9800 ft) and
live almost completely buried in the ground except during the wet season
when they spring back to life bringing a burst of colour when in bloom.
Within South Africa are to be found a variety of climates and diverse
topography ranging from areas with a winter rainy season to savannahs with
wet summers, from jungles to prairies, from lowlands to high-altitude plains
and finally from snowy peaks to mountain forests.
On the mountains north of Cape Town we encounter a great diversity of
genera and species including Aloe, Cotyledon, Pelargonium, Crassula,
Othonna, Haworthia, Euphorbia, Stapelia, Gasteria and Conophytum.
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